Polystichum kruckebergii
holly fern, Kruckeberg's sword fern fern

Distribution: Occurring in the Olympics and Cascades mountains in Washington; southeast Alaska to California, east to Montana, Idaho, Nevada and Utah.

Habitat: Open slopes, typically among rocks; tolerant of serpentine.

Origin: Native

Growth Duration: Perennial

Conservation Status: Not of concern


Terrestrial perennials with ascending stems.


Leaves monomorphic, erect, 1-2.5 dm, lacking bulblets; petiole 1/5 length of leaf or less, with sparse covering of light brown scales that become smaller distally; blade linear, 1-pinnate-pinnafitid, base narrowed; pinnae rhombic-ovate to short-falcate, proximal pinnae somewhat triangular, pinnae overlapping and twisted slightly out of blade plane, 0.5-1.5 cm, base oblique, acroscopic auricle distinct, margins shallowly cut to barely dentate or finely serrated, teeth spreading and spiny at tip, apex acute; microscales lanceolate, confined to costa on bottom surface.


Indusia entire; spores dark brown.

Accepted Name:
Polystichum kruckebergii W.H. Wagner
Publication: Amer. Fern J. 56: 4. 1966.

Synonyms & Misapplications:
(none provided)
Additional Resources:

PNW Herbaria: Specimen records of Polystichum kruckebergii in the Consortium of Pacific Northwest Herbaria database

WA Flora Checklist: Polystichum kruckebergii checklist entry

OregonFlora: Polystichum kruckebergii information

E-Flora BC: Polystichum kruckebergii atlas page

CalPhotos: Polystichum kruckebergii photos

16 photographs:
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