A Checklist of the Vascular Plants of Washington State
Hosted by the University of Washington Herbarium, Burke Museum
Hosted by the University of Washington Herbarium, Burke Museum
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Dryopteridaceae » Dryopteris carthusiana
Last updated 10/26/2019 by David Giblin.
Dryopteris carthusiana (Vill.) H.P. Fuchs[FNA2, HC2]
spinulose wood-fern, toothed wood-fern
Publication: Bull. Soc. Bot. France. 105: 339. 1959.
Origin: Native
selected vouchers: WTU
Notes: D. austriaca is European.
FNA2: "Dryopteris carthusiana is tetraploid. Dryopteris intermedia is one parent, as indicated by chromosome pairing in their hybrid D . × triploidea Wherry. The other parent is the hypothetical missing ancestral species " D . semicristata " (see discussion for D . cristata ). Dryopteris carthusiana hybridizes with five species; hybrids can be separated from D . intermedia by the lack of glandular hairs and by having 2-pinnate leaves."
References: (none)
Synonyms & Misapplied Names:
Dryopteris austriaca (Jacq.) Schinz & Thell. var. spinulosa (O.F. Muell.) Fiori
Dryopteris spinulosa (O.F. Muell.) Watt
Polypodium carthusianum Vill.
Polypodium spinulosum O.F. Muell.
