Checklist » Boraginaceae
Boraginaceae[Draft FNA, HC, HC2](Borage Family)
Synonyms: (none)

Notes: Taxonomy follows recent molecular studies:

Weigend, M., Federico Luebert, Marc Gottschling, Thomas L.P., Couvreur, Hartmut H. Hilger and James S. Miller. 2013. From capsules to nutlets—phylogenetic relationships in the Boraginales. Cladistics (2013) 1–11.

Nazaire, M., and L Hufford. 2012. A Broad Phylogenetic Analysis of Boraginaceae: Implications for the Relationships of Mertensia. Systematic Botany 37(3): pp. 758–783.

Family information last updated 2/9/2016 by David Giblin.

Taxon (native taxa in boldface)
Common Names
# Synonyms
Adelinia grandis (Douglas ex Lehm.) J.I. Cohen[HC2]
Adeline's hound's-tongue, Pacific hound's-tongue, grand hound
1 synonym
Amsinckia intermedia Fisch. & C.A. Mey.[HC, HC2]
fireweed fiddleneck, rancher's fiddleneck
2 synonyms
Amsinckia lycopsoides Lehm.[HC, HC2]
bugloss fiddleneck, tarweed fiddleneck
0 synonyms
Amsinckia menziesii (Lehm.) A. Nelson & J.F. Macbr.[HC, HC2]
harvest fiddleneck, rigid fiddleneck, rancher's fireweed
2 synonyms
Amsinckia retrorsa Suksd.[HC, HC2]
harvest fiddleneck, rigid fiddleneck
0 synonyms
Amsinckia spectabilis Fisch. & C.A. Mey.[HC, HC2]
seaside amsinckia, seaside fiddleneck
1 synonym
var. spectabilis[HC2, JPM]
seaside amsinckia, seaside fiddleneck
0 synonyms
Amsinckia tessellata A. Gray[HC, HC2]
bristly amsinckia, bristly fiddleneck, tessellate fiddleneck
0 synonyms
var. tessellata[HC2, JPM]
bristly amsinckia, bristly fiddleneck, tessellate fiddleneck
0 synonyms
Anchusa azurea Mill.[HC, HC2]
Italian alkanet, Italian bugloss
0 synonyms
var. azurea[HC2]
Italian alkanet, Italian bugloss
0 synonyms
Anchusa officinalis L.[HC, HC2]
common alkanet, common bugloss
0 synonyms
Andersonglossum occidentale (A. Gray) J.I. Cohen[HC2]
western hound's-tongue
1 synonym
Andersonglossum virginianum (L.) J.I. Cohen[HC2]
northern hound's-tongue
0 synonyms
Asperugo procumbens L.[HC, HC2]
catchweed, madwort
0 synonyms
Borago officinalis L.[HC, HC2]
borage, common borage
0 synonyms
Buglossoides arvensis (L.) I.M. Johnst.[HC2]
corn gromwell, field gromwell
1 synonym
Cryptantha affinis (A. Gray) Greene[HC, HC2]
quill cat's-eye, common cryptantha, slender cryptantha
1 synonym
Cryptantha ambigua (A. Gray) Greene[HC, HC2]
basin cat's-eye, obscure cryptantha, wilke's cryptantha
0 synonyms
Cryptantha echinella Greene[HC, HC2]
prickly cryptantha
0 synonyms
Cryptantha fendleri (A. Gray) Greene[HC, HC2]
sand dune cat's-eye, Fendler's cryptantha
0 synonyms
Cryptantha flaccida (Douglas ex Lehm.) Greene[HC, HC2]
weakstem cat's-eye, flaccid cryptantha
0 synonyms
Cryptantha gracilis Osterh.[HC2, JPM]
narrow-stem cryptantha, slender cryptantha
0 synonyms
Cryptantha grandiflora Rydb.[Draft FNA, HC2]
large-flowered cryptantha
1 synonym
Cryptantha intermedia (A. Gray) Greene[HC, HC2]
Clearwater cat's-eye, common cryptantha
1 synonym
var. hendersonii (A. Nelson) Jeps. & Hoover[HC, HC2]
Henderson's common cryptantha
1 synonym
Cryptantha kelseyana Greene[HC, HC2]
Kelsey's cryptantha
0 synonyms
Cryptantha minima Rydb.[HC2]
bracteate cryptantha
0 synonyms
Cryptantha pterocarya (Torr.) Greene[HC, HC2, JPM]
wingnut cryptantha
0 synonyms
var. pterocarya[HC2]
wingnut cryptantha
0 synonyms
Cryptantha rostellata (Greene) Greene[HC, HC2]
beaked cryptantha
1 synonym
Cryptantha scoparia A. Nelson[HC, HC2]
pinyon desert cat's-eye, desert cryptantha
0 synonyms
Cryptantha simulans Greene[HC, HC2]
pine woods cat's-eye, pine woods cryptantha
0 synonyms
Cryptantha torreyana (A. Gray) Greene[HC2, JPM2]
Torrey's cryptantha
1 synonym
var. torreyana[HC2]
Torrey's cryptantha
0 synonyms
Cryptantha watsonii (A. Gray) Greene[HC, HC2]
Watson's cryptantha
0 synonyms
Cynoglossum officinale L.[HC, HC2]
common hound's-tongue
0 synonyms
Dasynotus daubenmirei I.M. Johnst.[HC, HC2]
 
0 synonyms
Echium plantagineum L.[HC2]
salvation Jane, purple viper's bugloss
0 synonyms
Echium vulgare L.[HC, HC2]
common viper's bugloss
0 synonyms
Eritrichium argenteum W. Wight[Draft FNA, HC2]
alpine forget-me-not
3 synonyms
Eritrichium howardii (A. Gray) Rydb.[HC, HC2]
Howard's alpine forget-me-not
0 synonyms
Greeneocharis circumscissa (Hook. & Arn.) Rydb.[HC2]
cushion cryptantha, matted cryptantha
1 synonym
var. circumscissa[HC2]
 
3 synonyms
Gruvelia pusilla A. DC.[Draft FNA, HC2]
little gruvelia
1 synonym
Hackelia americana (A. Gray) Fernald[HC2]
nodding stickseed
3 synonyms
Hackelia californica (A. Gray) I.M. Johnst.[HC, HC2]
California stickseed
0 synonyms
Hackelia ciliata (Douglas ex Lehm.) I.M. Johnst.[HC, HC2]
Okanogan stickseed
0 synonyms
Hackelia cinerea (Piper) I.M. Johnst.[HC, HC2]
gray stickseed
0 synonyms
Hackelia cronquistii J.L. Gentry[HC, HC2]
Cronquist's stickseed
0 synonyms
Hackelia cusickii (Piper) Brand[HC, HC2]
Cusick's stickseed
0 synonyms
Hackelia davisii Cronquist[HC, HC2]
Davis's stickseed
0 synonyms
Hackelia diffusa (Lehm.) I.M. Johnst.[HC, HC2]
diffuse stickseed
0 synonyms
var. arida (Piper) R.L. Carr[HC2]
sagebrush stickseed, steppe stickseed
1 synonym
var. cottonii (Piper) R.L. Carr[HC2]
Cotton's stickseed
0 synonyms
var. diffusa[HC2]
diffuse stickseed
1 synonym
Hackelia floribunda (Lehm.) I.M. Johnst.[HC, HC2]
many-flowered stickseed
0 synonyms
Hackelia hispida (A. Gray) I.M. Johnst.[HC, HC2]
hispid stickseed, rough stickseed
0 synonyms
var. disjuncta R.L. Carr[HC2]
sagebrush stickseed
0 synonyms
var. hispida[HC2]
rough stickseed
0 synonyms
Hackelia micrantha (Eastw.) J.L. Gentry[HC, HC2]
meadow forget-me-not, blue stickseed
1 synonym
Hackelia patens (Nutt.) I.M. Johnst.[HC, HC2]
spreading stickseed
0 synonyms
var. patens[HC, HC2]
 
0 synonyms
Hackelia taylorii Harrod, Malmquist & R.L. Carr[HC2]
Taylor's stickseed
0 synonyms
Hackelia venusta (Piper) H. St. John[HC, HC2]
lesser showy stickseed
0 synonyms
Lappula desertorum Greene[HC2]
desert stickseed
0 synonyms
Lappula fremontii (Torr.) Greene[HC2]
Fremont's stickseed
0 synonyms
var. fremontii[HC2]
 
0 synonyms
Lappula longispina[HC2]
long-spined stickseed
0 synonyms
Lappula montana Greene[HC2]
montane stickseed
0 synonyms
Lappula occidentalis (S. Watson) Greene[HC2]
western stickseed
0 synonyms
Lappula squarrosa (Retz.) Dumort.[HC2, JPM2]
bristly sheepburr, bristly stickseed, common stickseed, European stickseed, bristly-fruited tickweed
1 synonym
Lithospermum incisum Lehm.[HC, HC2]
fringed stoneseed
0 synonyms
Lithospermum occidentale (Michx.) Muhl.[HC2]
 
1 synonym
Lithospermum ruderale Douglas ex Lehm.[HC, HC2]
 
1 synonym
Lycopsis arvensis L.[HC2]
annual bugloss, European bugloss, small bugloss
1 synonym
Mertensia ambigua Piper[HC2]
 
0 synonyms
Mertensia amoena A. Nelson[HC2]
beautiful bluebells, sagebrush bluebells
1 synonym
Mertensia bella Piper[HC, HC2]
beautiful bluebells, Oregon bluebells, lungwort
0 synonyms
Mertensia brachycalyx Piper[HC2]
short-sepaled bluebells
0 synonyms
Mertensia campanulata A. Nelson[HC, HC2]
Idaho bluebells
0 synonyms
Mertensia ciliata (E. James ex Torr.) G. Don[HC, HC2]
mountain bluebells, streamside bluebells, tall fringed bluebells
1 synonym
Mertensia coriacea A. Nelson[HC2]
green bluebells, leathery bluebells
0 synonyms
Mertensia cusickii Piper[HC2]
Cusick's bluebells
0 synonyms
Mertensia foliosa A. Nelson[HC2]
leafy bluebells
0 synonyms
Mertensia longiflora Greene[HC, HC2]
long-flowered bluebells, trumpet bluebells, long-flowered lungwort
0 synonyms
Mertensia oblongifolia (Nutt.) G. Don[HC, HC2]
oblong-leaved bluebell, sagebrush bluebell
3 synonyms
Mertensia paniculata (Aiton) G. Don[HC, HC2]
northern bluebell, tall bluebell
0 synonyms
var. borealis (J.F. Macbr.) L.O. Williams[HC, HC2, JPM]
northern bluebell
0 synonyms
var. paniculata[HC, HC2, JPM]
tall bluebell
0 synonyms
Mertensia platyphylla A. Heller[HC, HC2]
broadleaf bluebell
2 synonyms
Mertensia tweedyi Rydb.[HC2]
alpine bluebells
1 synonym
Mertensia umbratilis Greenm.[HC, HC2]
shade bluebell
0 synonyms
Myosotis alpestris F.W. Schmidt[Davis 1952, HC2]
Asian forget-me-not
2 synonyms
Myosotis arvensis (L.) Hill[HC, HC2]
field forget-me-not, field scorpion-grass
0 synonyms
Myosotis discolor Pers.[HC, HC2]
yellow and blue forget-me-not, yellow and blue scorpiongrass
1 synonym
Myosotis latifolia Poir.[HC2]
broad-leaved forget-me-not
0 synonyms
Myosotis laxa Lehm.[HC, HC2]
bay forget-me-not, small forget-me-not
0 synonyms
Myosotis micrantha Pall. ex Lehm.[HC, HC2]
blue scorpion-grass
0 synonyms
Myosotis scorpioides L.[HC, HC2]
common forget-me-not, true forget-me-not, water forget-me-not
1 synonym
Myosotis sylvatica Ehrh. ex Hoffm.[HC, HC2, Stace 1997]
woodland forget-me-not
1 synonym
Myosotis verna Nutt.[HC, HC2]
early forget-me-not, spring forget-me-not, early scorpion-grass, spring scorpion-grass
2 synonyms
Oreocarya glomerata (Pursh) Greene
cockscomb oreocarya
4 synonyms
Oreocarya humilis (A. Gray) Greene
 
1 synonym
Oreocarya hypsophila (I. M. Johnston) Hasenstab & M.G. Simpson
 
1 synonym
Oreocarya leucophaea (Douglas ex Lehm.) Greene
gray oreocarya
1 synonym
Oreocarya montana R.B. Kelley & Ripma
 
0 synonyms
Oreocarya propria A. Nelson & J.F. Macbr.
 
1 synonym
Oreocarya salmonensis A. Nelson & J.F. Macbr.
 
1 synonym
Oreocarya sobolifera (Payson) R.B. Kelley
 
1 synonym
Oreocarya spiculifera Piper[FNA, OFP]
Snake River cat's-eye, Snake River oreocarya
2 synonyms
Oreocarya subretusa (I.M. Johnst.) Abrams
 
0 synonyms
Oreocarya thompsonii (I.M. Johnst.) Abrams
Thompson's cat's-eye, Thompson's oreocarya
1 synonym
Pectocarya penicillata (Hook. & Arn.) A. DC.[HC2, IFBC]
shortleaf combseed, winged pectocarya
1 synonym
Pectocarya setosa A. Gray[HC, HC2]
bristly combseed
0 synonyms
Pentaglottis sempervirens (L.) Tausch ex L.H. Bailey[HC2, IFBC]
green alkanet
1 synonym
Plagiobothrys bracteatus (Howell) I.M. Johnst.[HC2]
 
0 synonyms
Plagiobothrys cognatus (Greene) I.M. Johnst.[HC2, JPM]
cognate popcorn-flower
1 synonym
Plagiobothrys cusickii (Greene) I.M. Johnst.[HC2, JPM]
matted popcorn-flower
3 synonyms
Plagiobothrys figuratus (Piper) I.M. Johnst. ex M. Peck[HC, HC2]
fragrant popcorn-flower
1 synonym
var. figuratus[HC2]
fragrant popcorn-flower
2 synonyms
Plagiobothrys hirtus (Greene) I.M. Johnst.
 
0 synonyms
Plagiobothrys hispidulus (Greene) I.M. Johnst.[HC2, JPM]
harsh popcorn-flower
2 synonyms
Plagiobothrys kingii (S. Watson) A. Gray[HC2]
Harkness's plagiobothrys
0 synonyms
var. harknessii (Greene) Jeps.[HC2]
 
1 synonym
Plagiobothrys leptocladus (Greene) I.M. Johnst.[HC, HC2]
alkali popcorn-flower, slender-branched popcorn-flower
0 synonyms
Plagiobothrys mollis (A. Gray) I.M. Johnst.[HC, HC2]
soft plagiobothrys
0 synonyms
var. mollis[HC2]
 
0 synonyms
Plagiobothrys nothofulvus (A. Gray) A. Gray[HC, HC2]
rusty popcorn-flower
0 synonyms
Plagiobothrys salsus (Brandegee) I.M. Johnst.[HC2]
 
0 synonyms
Plagiobothrys scouleri (Hook. & Arn.) I.M. Johnst.[HC, HC2]
Scouler's popcorn-flower
4 synonyms
Plagiobothrys stipitatus (Greene) I.M. Johnst.[HC2, JPM]
stipitate popcorn flower
0 synonyms
var. micranthus (Piper) I.M. Johnst.[HC2]
 
0 synonyms
Plagiobothrys tenellus (Nutt. ex Hook.) A. Gray[HC, HC2]
Pacific popcorn-flower, slender popcorn-flower
1 synonym
Pulmonaria ovalis Bast.
oval lungwort
1 synonym
Sonnea hispida (A. Gray) Greene[HC2]
bristly sonnea
0 synonyms
Symphytum asperum Lepech.[HC, HC2]
prickly comfrey, rough comfrey
1 synonym
Symphytum grandiflorum DC.[Stace 1997]
creeping comfrey
1 synonym
Symphytum officinale L.[HC, HC2]
common comfrey
0 synonyms
Symphytum orientale L.[Stace 1997]
white comfrey
0 synonyms
Symphytum ×uplandicum Nyman[HC2, Stace 1997]
hybrid comfrey
0 synonyms
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